Component | Descriptor name | Formula | Expected properties | Reference |
---|---|---|---|---|
Richness | Species density | Standardize species richness per unit area | ||
Margelef | Adjusted species richness by |
Margelef (1958) | ||
Evenness | Pielou index | Evenness based on the Shannon-Wiener index |
||
Heip | Sensitive to rare species | |||
Heterogeneity | Shannon-Wiener | Sensitive to rare species | ||
Simpson diversity | Sensitive to dominant species | |||
Taxonomy | Average taxonomic distinctness | natural extensions taxonomic of Simpson relatedness diversity including | ||
Variation in taxonomic distinctness | Evenness of the taxonomic level distribution in the taxonomic tree |
Location | Latitude | Longitude | Water temperature | Salinity | Water depth | |||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
Spring | Summer | Autumn | Winter | Spring | Summer | Autumn | Winter | Spring | Summer | Autumn | Winter | |||
1 | 26.189 | 119.652 | 20.8 | 26.2 | 21.9 | 14.9 | 28.0 | 29.5 | 29.1 | 27.5 | 11.2 | 10.5 | 10.7 | 10.9 |
2 | 26.240 | 119.741 | 21.0 | 27.1 | 21.9 | 14.7 | 30.3 | 32.5 | 29.4 | 31.3 | 10.2 | 13.6 | 13.2 | 11.7 |
3 | 26.288 | 119.842 | 20.4 | 26.2 | 21.5 | 15.3 | 32.2 | 33.0 | 28.2 | 33.1 | 20.8 | 18.2 | 19.9 | 19.6 |
4 | 26.111 | 119.669 | 21.7 | 26.5 | 22.6 | 15.5 | 25.8 | 29.6 | 25.6 | 27.0 | 10.6 | 11.6 | 8.2 | 9.1 |
5 | 26.121 | 119.770 | 20.9 | 27.4 | 21.8 | 16.2 | 30.3 | 31.6 | 28.9 | 32.2 | 12.7 | 11.5 | 10.0 | 13.1 |
6 | 26.131 | 119.867 | 20.9 | 25.6 | 22.7 | 15.6 | 23.8 | 32.5 | 29.0 | 34.0 | 19.7 | 23.4 | 22.3 | 22.6 |
7 | 26.040 | 119.697 | 21.1 | 27.0 | 22.6 | 14.8 | 23.7 | 25.9 | 19.8 | 27.1 | 7.3 | 7.1 | 8.2 | 9.1 |
8 | 26.018 | 119.780 | 21.2 | 27.0 | 22.7 | 14.7 | 28.5 | 31.1 | 16.1 | 32.3 | 13.1 | 12.7 | 14.8 | 15.1 |
9 | 25.987 | 119.863 | 22.2 | 26.6 | 22.5 | 13.3 | 20.7 | 31.5 | 30.0 | 36.2 | 18.3 | 22.0 | 19.0 | 17.8 |
10 | 25.931 | 119.770 | 22.1 | 27.7 | 22.5 | 13.3 | 25.5 | 31.3 | 23.7 | 31.0 | 15.7 | 16.6 | 13.7 | 14.3 |
11 | 25.850 | 119.833 | 21.1 | 26.8 | 22.6 | 13.0 | 30.8 | 34.3 | 25.3 | 33.0 | 25.6 | 24.7 | 24.3 | 24.7 |
Order | Family | Genus | Species | Spring | Summer | Autumn | Winter | ||||
---|---|---|---|---|---|---|---|---|---|---|---|
abundance (ind.) | biomass (g) | abundance (ind.) | biomass (g) | abundance (ind.) | biomass (g) | abundance (ind.) | biomass (g) | ||||
Carcharhiniformes | Carcharhinidae | -- | -- | 7 | 409.6 | -- | -- | -- | -- | ||
Rajiformes | Rhinobatidae | -- | -- | -- | -- | -- | -- | 2 | 886.2 | ||
-- | -- | -- | -- | 2 | 693.3 | -- | -- | ||||
1 | 300.0 | 3 | 455.8 | -- | -- | 1 | 351.7 | ||||
Myliobatiformes | Dasyatidae | -- | -- | -- | -- | -- | -- | 1 | 210.5 | ||
1 | 293.0 | 1 | 1468.0 | -- | -- | -- | -- | ||||
1 | 641.0 | 4 | 1478.5 | 7 | 1987.5 | -- | -- | ||||
1 | 10018.0 | -- | -- | 2 | 4339.3 | -- | -- | ||||
1 | 299.9 | -- | -- | -- | -- | -- | -- | ||||
Anguilliformes | Muraenesocidae | 26 | 1120.4 | 52 | 2951.4 | 2 | 245.7 | 7 | 226.5 | ||
Muraenidae | -- | -- | 2 | 124.6 | -- | -- | -- | -- | |||
Congridae | -- | -- | -- | -- | -- | -- | 1 | 30.0 | |||
Ophichthidae | 9 | 211.8 | 4 | 151.2 | 11 | 214.3 | -- | -- | |||
-- | -- | 6 | 131.4 | -- | -- | -- | -- | ||||
-- | -- | 31 | 707.5 | -- | -- | -- | -- | ||||
-- | -- | 2 | 61.0 | -- | -- | -- | -- | ||||
Lophiiformes | Antennariidae | -- | -- | 2 | 51.2 | -- | -- | -- | -- | ||
Gonorynchiformes | Gonorynchidae | -- | -- | 5 | 140.5 | -- | -- | -- | -- | ||
Siluriformes | Bagridae | 2 | 238.8 | -- | -- | 5 | 109.3 | 3 | 59.01 | ||
Clupeiformes | Clupeidae | 49 | 1050.6 | -- | -- | 1 | 88.3 | 6 | 158.9 | ||
20 | 312.8 | 1 | 4.5 | -- | -- | -- | -- | ||||
Pristigasteridae | 7 | 160.5 | 1 | 13.4 | 77 | 1913.2 | 4 | 150.7 | |||
Engraulidae | 110 | 1944.8 | 182 | 2829.6 | 90 | 866.2 | 1 | 3.3 | |||
211 | 2422.7 | 19 | 231.5 | 546 | 5261.7 | 1250 | 9977.8 | ||||
330 | 1020.0 | 13 | 30.5 | 255 | 2002.1 | 20 | 44.3 | ||||
7 | 72.7 | 5 | 13.5 | 15 | 147.2 | -- | -- | ||||
2 | 20.1 | -- | -- | -- | -- | 24 | 170.7 | ||||
-- | -- | -- | -- | 3 | 22.2 | -- | -- | ||||
99 | 102.4 | 8 | 21.1 | 8 | 7.3 | -- | -- | ||||
30 | 52.2 | -- | -- | -- | -- | -- | -- | ||||
Aulopiformes | Synodontidae | 1 | 3.9 | -- | -- | -- | -- | -- | -- | ||
-- | -- | 5 | 6.5 | -- | -- | -- | -- | ||||
63 | 1505.8 | 1144 | 34623.2 | 2284 | 29280.9 | 279 | 9622.1 | ||||
5 | 38.9 | -- | -- | -- | -- | -- | -- | ||||
4 | 258.2 | 56 | 2738.0 | 1 | 32.4 | 28 | 1612.3 | ||||
Scorpaeniformes | Scorpaenidae | -- | -- | 1 | 1.7 | -- | -- | -- | -- | ||
-- | -- | -- | -- | -- | -- | 10 | 157.2 | ||||
Sebastidae | 2 | 1.8 | 4 | 30.2 | 2 | 152 | -- | -- | |||
Synanceiidae | -- | -- | -- | -- | 1 | 1.0 | -- | -- | |||
Platycephalidae | -- | -- | 7 | 61.4 | -- | -- | -- | -- | |||
5 | 208.4 | 2 | 108.2 | 4 | 42.8 | 1 | 13.1 | ||||
2 | 169.9 | 4 | 52.1 | 1 | 85.2 | -- | -- | ||||
Triglidae | 328 | 2254.7 | 3 | 82.25 | -- | -- | 3 | 258.7 | |||
Mugiliformes | Mugilidae | 1 | 28.3 | -- | -- | 2 | 67.4 | -- | -- | ||
-- | -- | -- | -- | -- | -- | 2 | 35.0 | ||||
1 | 26.9 | 3 | 80.1 | -- | -- | -- | -- | ||||
Syngnathiformes | Syngnathidae | -- | -- | 1 | 0.9 | -- | -- | -- | -- | ||
Fistulariidae | -- | -- | 5 | 13.5 | -- | -- | -- | -- | |||
Perciformes | Lateolabracidae | -- | -- | -- | -- | 8 | 4273.0 | 5 | 1063.0 | ||
Leiognathidae | -- | -- | -- | -- | -- | -- | 1 | 7.04 | |||
-- | -- | 3 | 1.3 | -- | -- | -- | -- | ||||
-- | -- | -- | -- | 1 | 9.9 | -- | -- | ||||
6 | 79.37 | -- | -- | -- | -- | 3 | 22.15 | ||||
982 | 5120.7 | 1654 | 10399.9 | 299 | 1545.3 | 107 | 282.69 | ||||
Terapontidae | -- | -- | 1 | 11.1 | 2 | 30.4 | -- | -- | |||
Siganidae | -- | -- | 4 | 35.35 | -- | -- | -- | -- | |||
-- | -- | 168 | 1824.5 | -- | -- | -- | -- | ||||
Carangidae | -- | -- | 10 | 340.8 | -- | -- | -- | -- | |||
-- | -- | 141 | 2661.9 | -- | -- | -- | -- | ||||
2647 | 6918.6 | -- | -- | -- | -- | -- | -- | ||||
Sciaenidae | 1 | 32.1 | 6182 | 20174.7 | 52 | 1345.5 | -- | -- | |||
-- | -- | -- | -- | -- | -- | 7 | 31.34 | ||||
-- | -- | 3 | 71.2 | -- | -- | -- | -- | ||||
6 | 465.3 | -- | -- | 21 | 643.9 | 5 | 405.2 | ||||
17 | 1168.3 | 353 | 8112.0 | 3 | 250.8 | -- | -- | ||||
1 | 100.7 | -- | -- | -- | -- | -- | -- | ||||
1 | 51.9 | 2 | 199.2 | -- | -- | -- | -- | ||||
6 | 149.3 | 81 | 4839.3 | 54 | 1116.6 | -- | -- | ||||
130 | 3563.6 | 38 | 334.0 | 228 | 4161.3 | 544 | 12549.7 | ||||
1 | 71.6 | -- | -- | -- | -- | -- | -- | ||||
Sparidae | -- | -- | -- | -- | -- | -- | 1 | 428.5 | |||
357 | 1268.6 | 122 | 1601.0 | -- | -- | -- | -- | ||||
1 | 218.9 | -- | -- | -- | -- | -- | -- | ||||
-- | -- | 6 | 401.8 | -- | -- | -- | -- | ||||
Priacanthidae | -- | -- | 156 | 2488.4 | -- | -- | -- | -- | |||
Apogonidae | -- | -- | -- | -- | 4 | 7.3 | -- | -- | |||
1 | 15.0 | -- | -- | 4 | 8.2 | -- | -- | ||||
Hapalogenyidae | -- | -- | 1 | 3.8 | -- | -- | -- | -- | |||
1 | 11.0 | -- | -- | -- | -- | -- | -- | ||||
Callionymidae | -- | -- | 17 | 148.0 | -- | -- | -- | -- | |||
-- | -- | 3 | 21.5 | -- | -- | -- | -- | ||||
Uranoscopidae | -- | -- | 3 | 105.5 | -- | -- | -- | -- | |||
-- | -- | -- | -- | 1 | 47.9 | -- | -- | ||||
Mullidae | -- | -- | 1524 | 15023.7 | -- | -- | 1 | 17.3 | |||
Sphyraenidae | -- | -- | 6 | 203.4 | -- | -- | -- | -- | |||
Sillaginidae | 2 | 116.1 | 31 | 175.7 | 4 | 149.2 | 26 | 854.8 | |||
Stromateidae | -- | -- | 4 | 190.0 | -- | -- | -- | -- | |||
-- | -- | 10 | 500.6 | -- | -- | -- | -- | ||||
303 | 2227.8 | 24 | 2412.7 | 15 | 2391.2 | 3 | 45.5 | ||||
-- | -- | 25 | 1108.1 | -- | -- | -- | -- | ||||
Centrolophidae | 59 | 405.1 | 124 | 4162.3 | -- | -- | -- | -- | |||
Trichiuridae | -- | -- | 8 | 468.5 | 2 | 21.5 | -- | -- | |||
1 | 31.6 | 151 | 7776.8 | 20 | 488.2 | 9 | 356.3 | ||||
Polynemidae | -- | -- | 13485 | 30886.6 | 1929 | 13965.1 | -- | -- | |||
-- | -- | -- | -- | 3 | 234.5 | -- | -- | ||||
Scombridae | -- | -- | -- | -- | 3 | 2056.1 | 6 | 3987.0 | |||
16 | 106.3 | -- | -- | -- | -- | -- | -- | ||||
Gobiidae | 3 | 33.6 | -- | -- | -- | -- | -- | -- | |||
448 | 2452.1 | 70 | 149.7 | 436 | 1899.4 | 160 | 1169.6 | ||||
2 | 25.8 | -- | -- | -- | -- | 2 | 29.7 | ||||
9 | 108.4 | 60 | 593.9 | 8 | 80.2 | 81 | 840.3 | ||||
67 | 1081.7 | 12 | 40.8 | 5 | 26.8 | 1 | 4.5 | ||||
3 | 13.4 | -- | -- | -- | -- | -- | -- | ||||
2 | 13.0 | -- | -- | -- | -- | -- | -- | ||||
-- | -- | -- | -- | -- | -- | 2 | 29.7 | ||||
Pleuronectiformes | Paralichthyidae | -- | -- | 15 | 661.3 | -- | -- | -- | -- | ||
1 | 144.9 | -- | -- | -- | -- | -- | -- | ||||
1 | 15.7 | -- | -- | -- | -- | -- | -- | ||||
Pleuronectidae | 1 | 4.2 | -- | -- | -- | -- | -- | -- | |||
Cynoglossidae | -- | -- | 7 | 71.5 | -- | -- | -- | -- | |||
233 | 8567.4 | 352 | 4290.4 | 145 | 2820.1 | 276 | 6396.6 | ||||
-- | -- | -- | -- | -- | -- | 1 | 23.3 | ||||
2 | 29.0 | -- | -- | -- | -- | 10 | 227.7 | ||||
1 | 9.9 | -- | -- | -- | -- | 1 | 4.3 | ||||
8 | 450.3 | 35 | 1899.2 | 6 | 500.5 | 9 | 631.03 | ||||
Soleidae | -- | -- | 19 | 191.1 | -- | -- | -- | -- | |||
Monacanthidae | -- | -- | 78.6 | 328.3 | -- | -- | -- | -- | |||
-- | -- | 5 | 33.5 | -- | -- | -- | -- | ||||
Tetraodontidae | 4 | 118.6 | 4 | 23.9 | -- | -- | 5 | 132.6 | |||
-- | -- | -- | -- | 2 | 38.7 | -- | -- | ||||
7 | 195.4 | 352 | 3805.1 | 65 | 2333.0 | 11 | 252.1 | ||||
-- | -- | -- | -- | -- | -- | 2 | 280.7 | ||||
-- | -- | 1 | 9.9 | -- | -- | -- | -- | ||||
-- | -- | -- | -- | 5 | 1296.0 | -- | -- | ||||
-- | -- | 12 | 397.5 | -- | -- | -- | -- | ||||
-- | -- | -- | -- | -- | -- | 3 | 430.79 | ||||
-- | -- | 616 | 13003.2 | 22 | 1714.7 | -- | -- |
Species | Season | ||||||||
---|---|---|---|---|---|---|---|---|---|
spring | 1.11 | 0.26 | 1.46 | 99.93 | 25.54 | 47.18 | |||
summer | 33.63 | 54.28 | 61.96 | 21.79 | 67.94 | 13.75 | 20.40 | ||
autumn | 2.01 | 2.39 | 59.23 | 15.82 | 5.85 | 1.95 | |||
winter | 33.46 | 34.99 | 21.90 | 24.35 | 23.56 | ||||
spring | 87.28 | 90.83 | 0.73 | 12.24 | 19.59 | ||||
summer | 13.26 | 41.35 | 94.70 | 17.54 | 4.63 | 5.07 | |||
autumn | 8.39 | 53.11 | 19.31 | 19.24 | 23.54 | ||||
winter | 37.48 | 80.08 | 3.39 | 2.12 | |||||
summer | 29.78 | 0.91 | 51.16 | 59.84 | 19.48 | ||||
autumn | 0.47 | 3.06 | 37.21 | 37.00 | |||||
spring | 93.40 | 0.00 | 2.65 | 14.49 | |||||
summer | 32.16 | 14.34 | 8.87 | 15.31 | |||||
autumn | 11.99 | 0.00 | 1.20 | ||||||
winter | 54.67 | 1.63 | 0.86 | ||||||
spring | 0.92 | 16.79 | 10.90 | ||||||
summer | 14.95 | 0.27 | 0.12 | ||||||
autumn | 6.39 | 7.22 | |||||||
winter | 0.88 | 0.00 | |||||||
spring | 24.14 | 45.23 | |||||||
summer | 19.86 | 0.12 | |||||||
autumn | 21.41 | ||||||||
spring | 28.13 | ||||||||
summer | 18.21 | ||||||||
winter | 99.99 |
Spring | Summer | Autumn | Winter | ||
---|---|---|---|---|---|
Species richness | 64 | 78 | 49 | 46 | |
7.137 | 7.547 | 5.452 | 5.643 | ||
Evenness | 0.556 | 0.401 | 0.515 | 0.503 | |
0.144 | 0.061 | 0.134 | 0.123 | ||
Heterogeneity | 2.311 | 1.745 | 2.005 | 1.924 | |
1 − λ | 0.811 | 0.688 | 0.782 | 0.757 | |
Taxonomy | Δ+ | 78.556 | 58.159 | 74.553 | 73.279 |
˄+ | 96.900 | 84.564 | 95.342 | 96.832 |