Species | L | BD | EP | NR | NL | T | SL | GL |
|
|
|
|
SW% | GS% | D% | Country | Reference |
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
|
692–826 | 36–43 | 96–116 | 71–88 | 97–114 | 29–41 | 35–45 | 19–23 | 18.7* | 7.7** | 27.5** | 1.3** | 120–187 | 44–56 | 95–109 | Brazil |
|
|
842–1025 | 42–55 | 116–149 | 69–93 | 99–119 | 24–36 | 40–49 | 17–22 | 19.7* | 9.6** | 29.3** | 1.5** | 179–249 | 38–51 | 108–126 | Chile |
|
|
780–960 | 38–46 | 114–130 | 65–81 | 99–105 | 22–36 | 36–44 | 18–25 | 20.8* | 9.1* | 34.3* | 1.8* | 174 | 50 | 117 | Australia |
|
700–940 | 37–50 | 113–140 | 70–85 | 95–110 | 20–27 | 39–47 | 18–24 | – | – | – | – | – | – | – | Argentina | Agüera de Doucet and Doucet (1986) | |
689–880 | 38–46 | 78–123 | 55–90 | 92–124 | 21–32 | 34–48 | 17–26 | – | – | – | 1.2* | 147–256 | 41–49 | 68–106 | Australia |
|
|
782–927 | 92–120 | 103–139 | 58–76 | 84–105 | 28–37 | 51–53 | 17–26 | 6.6–8.5 | 8.5–10 | 23–32 | 1.4–2.2 | 194–282 | 37–57 | 108–157 | India |
|
|
805–1075 | 42–57 | 84–111 | 84–75 | 80–119 | 24–39 | 39–51 | 17–27 | 16–22 | 7.1–12 | 22–41 | 1.0–1.7 | 170–225 | 40–62 | 77–136 | India |
|
|
as |
1000–2000 | 42–70 | 145–170 | 64–82 | 103–120 | 28–49 | 42–49 | 20–26 | 16.7* | 8.3* | 14.3* | 1.4* | 198* | 62* | 92* | Argentina |
|
as |
1000–1200 | 32–60 | 125* | 125* | 113–131 | 29–36 | 42–52 | 22–27 | 19–35 | 8–11 | 28–38 | 1.3* | 185* | 51* | 95* | USA |
|
|
818–970 | 45–53 | 71–93 | 54–77 | 105–132 | 28–38 | 33–45 | 18–22 | 16–22 | 6.4–8.8 | 24–33 | 1.5** | 138–208 | 44–61 | 79** | Vietnam |
|
710–903 | 40–50 | 83–93 | 53–68 | 98–110 | 33–40 | 43–48 | 20–28 | 16–20 | 6.7–9.3 | 18–28 | – | 154–200 | 47–61 | 80–90 | India |
|
|
as |
737–870 | 37–44 | 68–93 | 72–83 | 90–120 | 20–30 | 32–45 | 17–23 | 20.7** | 8.3** | 32.3** | 1.2** | 133–198 | 42–59 | 74–99 | Thailand |
|
|
889–1192 | 51–73 | 130–157 | 81–108 | 116–143 | 32–45 | 40–49 | 22–27 | 15–23 | 7.2–10 | 22–34 | 1.3–2.3 | 153–208 | 48–59 | 102–120 | China |
|
|
699–876 | 33–40 | 86–91 | 62–78 | 97–106 | 29–34 | 41–47 | 17–19 | 26.6* | 8.8** | 27.4** | 1.4** | 170–220 | 36–47 | 90 | Ireland |
|
|
594–848 | 31–56 | 80–97 | 56–84 | 96–109 | 23–34 | 25–50 | 16–22 | 17.1* | 6.6** | 19.5** | 1.5* | 120–220 | 40–65 | 84–91 | Egypt | Abd–Elgawad and Ameen (2005) |
|
785–294 | 43–50 | 104–128 | 73–90 | 97–111 | 29–40 | 36–46 | 17–30 | 19.9* | 7.9** | 24.1** | 1.4** | 133–209 | 47–65 | 112 | USA |
|
|
721–913 | 43–55 | 101–145 | 72–93 | 100–122 | 29–41 | 41–49 | 20–28 | 16.5* | 7.7** | 26.1** | 1.4** | 150–200 | 51–64 | 100–122 | USA |
|
|
510–800 | 38–60 | 80–100 | 80–90 | – | – | – | – | – | – | – | – | – | – | – | Brazil |
|
|
554–837 | – | – | – | – | 30.9* | 43–60 | 26–30 | 18–22 | 5.9–8.2 | 18–37 | 1.1* | 167** | 55** | – | USA |
|
|
573–788 | 35–46 | 109–138 | 72–85 | 93–109 | 24–32 | 35–48 | 18–23 | 17.6* | 6.7** | 23.0** | 1.1** | 187 | 49 | 121 | India |
|
724–864 | 41–48 | 96–113 | 63–80 | 89–109 | 29–36 | 30–40 | 21–31 | 17–20 | 7.6–8.6 | 22–27 | 1.4–1.8 | 155–210 | 49–68 | 101–111 | India |
|
|
609–916 | 26–50 | 78–109 | 62–83 | 90–116 | 18–33 | 37–48 | 19–26 | 16–28 | 6.5–8.2 | 25–37 | 1.0–1.5 | 116–225 | 49–64 | 86–106 | India |
|
|
as |
840–950 | 40–48 | 92–100 | 80–88 | 104–112 | 28–36 | 44–48 | 20–24 | – | – | – | 2.9* | 170* | 47* | 84* | China |
|
as |
508–916 | 34–48 | 93–141 | 54–87 | 78–115 | 28–46 | 34–48 | 16–27 | – | – | – | – | 138–274 | 40–69 | 100–172 | Australia | Plichta et al. (2009) |
as |
864–1130 | 49–84 | 71–146 | 67–112 | 100–149 | 26–40 | 40–51 | 18–26 | – | – | – | – | – | – | – | USA |
|
as |
720–1013 | 38–43 | 112–133 | 80–110 | 100–105 | 30–42 | 35–42 | 20–22 | 18–24 | 7.2–9.8 | 19–25 | 1.4** | 144–191 | 48–65 | 110–126 | Pakistan |
|
|
805–1046 | 48–56 | 110–168 | 61–95 | 99–123 | 24–38 | 41–49 | 18–22 | 15.5* | 7.8** | 30.0** | 1.1** | 196 | 36–50 | 113** | USA |
|
960–1010 | 48–80 | 107–116 | 89–95 | 115–130 | 37–47 | 48–52 | 21–24 | – | – | – | – | – | – | – | USA |
|
|
as |
8000–1000 | 65–98 | 102–131 | 84–114 | 113–139 | 37–49 | 42–52 | 17–24 | – | – | – | – | – | – | – | USA |
|
|
800–1100 | 44–50 | 139–176 | 96–112 | 122–134 | 35–43 | 46–54 | 17–24 | 18–22 | 7–9 | 23–31 | 1.6* | 188 | 43 | 122 | USA |
|
|
614–801 | 38–47 | 108–145 | 61–83 | 89–108 | 21–36 | 30–47 | 18–32 | 21.7* | 6.8** | 27.6** | 1.1** | 130–196 | 43–70 | 114–149 | Mexico |
|
|
530–775 | 34–46 | 75–102 | 64–75 | 88–106 | 21–32 | 37–49 | 17–24 | 14–18 | 5.6–7.9 | 21–33 | 1.1–1.7 | 202–301 | 38–56 | 81–108 | S. Africa | Malan et al. (2014) |
|
970–1100 | 43–70 | – | – | 150–150 | 36–65 | 43–55 | 24–32 | 95–100 | 51–95 | 11–97 | – | – | – | – | USA |
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
777–1009 | 40–58 | 104–147 | 52–61 | 105–126 | 27–49 | 35–54 | 19–27 | 20.1* | 7.9** | 43.0** | 1.5* | 130–259 | 43–62 | 92–133 | S. Africa |
|
|
648–736 | 38–48 | 78–120 | 54–88 | 85–123 | 20–29 | 30–42 | 12–21 | 15.1* | 6.5** | 14.0** | 1.3** | 156 | 46 | 88 | Egypt |
|
as |
500–750 | 32–42 | 60–84 | 60–80 | 80–100 | 25–45 | 31–45 | 20–31 | – | – | – | – | 110–180 | 40–75 | 72–91 | Mexico |
|
|
848–1044 | 36–45 | 130–150 | – | 110–128 | 30–41 | 48–55 | 19–25 | – | – | – | 1.7* | 246 | 44 | 118 | N. Zealand |
|
as |
848–1044 | 36–45 | 130–150 | – | 110–128 | 30–41 | 48–55 | 19–25 | – | – | – | 1.7* | 246 | 44 | 118 | N. Zealand |
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
Species | L | BD | EP | NR | NL | T |
|
|
|
|
D% | E% | Country | Reference |
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
|
567-612 | 20-24 | 89-115 | 76-93 | 107-132 | 98-115 | 24-29 | 4.4-5.5 | 5.1-6.1 | 7.3* | 83-92 | 89–109 | Brazil | Andaló et al. (2006) |
|
578-666 | 19-26 | 101-126 | 79-101 | 124-144 | 94-107 | 25-31 | 4.8-5.7 | 5.7-7.1 | 5.7* | 79-94 | 149-182 | Chile |
|
|
512-671 | 18-31 | 87-110 | 72-93 | 100-139 | 83–112 | 17-30 | 4.0-5.1 | 5.7-7.0 | 6.0* | 76-92 | 103-130 | Australia |
|
530-660 | 22-30 | 93-108 | 80-90 | 110-130 | 84-105 | 23* | 4.4* | 5.7* | 4.8* | 81* | 106* | Argentina | Agüera de Doucet and Doucet (1986) | |
537-587 | 19-22 | 87-104 | 67-83 | 112-121 | 94-111 | 25-31 | 3.3-3.6 | 3.6-4.6 | 7.0* | 73-88 | 87-105 | Australia |
|
|
474-568 | 22-28 | 110-127 | 61-90 | 90-115 | 57-90 | 19-25 | 4.7-6.1 | 5.5-9.3 | 3.4-7.5 | 105-139 | 131-211 | India | Bhat et al. (2019a) | |
453-167 | 19-27 | 72-102 | 50-74 | 83-106 | 47-89 | 19-29 | 4.9-7.4 | 6.0-12 | 3.7-6.5 | 78–107 | 105-189 | India |
|
|
|
610-710 | 24-38 | 68-112 | 82-116 | 101-150 | 70-105 | 18.3* | 3.7* | 6.5* | 4.3* | 80* | 141* | Argentina |
|
|
619-671 | 23-29 | 112* | 108* | 130-139 | 104-112 | 22-28 | 4.6-5.4 | 5.8-6.3 | 6.0* | 83* | 97* | USA |
|
|
497-595 | 18-22 | 91–103 | 75-86 | 107-120 | 83-97 | 26-30 | 4.5-5.1 | 6-6.7 | 7.2* | 78-88 | 98–114 | Vietnam |
|
525-615 | 18-25 | 88-96 | 68-85 | 98-120 | 95-108 | 24-32 | 4.6-5.9 | 5.2-6.1 | – | 74-86 | 89-92 | India |
|
|
|
502-565 | 19-23 | 81-95 | 78-94 | 106-117 | 91-131 | 23-27 | 5-5 | 4-6 | 8.0* | 76-87 | 64–95 | Thailand |
|
|
566-687 | 21-25 | 100-122 | 85-106 | 118-146 | 86-111 | 24–29 | 4.2-4.9 | 5.9-6.8 | 6.0-7.4 | 80-93 | 103–121 | China |
|
|
588-692 | 15-22 | 96-128 | 96-105 | 126-141 | 62-74 | 29-42 | 4.4-5.3 | 8.5-10.5 | 4.4* | 76-96 | 160-180 | Ireland |
|
|
484-515 | 18-23 | 81-94 | 78-100 | 100-119 | 53-75 | 20-27 | 4.2-5.2 | 6.8-9.1 | 6.9* | 74-82 | 100-170 | Egypt | Abd-Elgawad and Ameen (2005) |
|
554-609 | 19-23 | 101-122 | 68-107 | 123-142 | 91-113 | 25-32 | 3.9-4.9 | 5.3-6.6 | 7.2* | 71–90 | 95–134 | USA |
|
|
547-651 | 17-26 | 97-113 | 74-94 | 110-139 | 86-108 | 23–34 | 4.1-5.3 | 5.5-6.9 | 6.8* | 70-93 | 106 | USA |
|
|
– | – | – | – | – | – | – | – | – | – | – | – | Brazil |
|
|
– | – | – | – | – | – | – | – | – | – | – | – | USA |
|
|
479-573 | 19-22 | 88-107 | 72-85 | 109-123 | 93-109 | 25-27 | 4.3-4.8 | 4.5-5.6 | – | 79-90 | 83-103 | India |
|
511-546 | 21-24 | 92-108 | 63–73 | 86-103 | 24–34 | 22-25 | 5-6 | 4.6-5.4 | 2.8-5.2 | 77-96 | 100-118 | India |
|
|
516-598 | 21-25 | 98-123 | 82-101 | 102-129 | 80-112 | 24-27 | 4.5-5.4 | 4.9-5.7 | 5.6-8.1 | 83-97 | 93–136 | India | Bhat et al. (2021b) | |
as |
528-632 | 20-24 | 104-116 | 96-104 | 120-136 | 68-80 | – | – | 6.6-8.6 | 6.3* | 81* | 150-180 | China |
|
as |
551-682 | 18-29 | 92-111 | 81-105 | 110-130 | 76-141 | 23-32 | 16-23 | 11-21 | 6.8* | 73-92 | 73-138 | Australia | Plichta et al.(2009) |
as |
506-631 | 21-28 | 116-175 | 79-103 | 115-181 | 82-108 | – | – | – | 6.0* | 77* | 88* | USA |
|
as |
558-624 | 19-23 | 95-106 | 73-90 | 113-125 | 95-110 | 25-29 | 4.7-5.3 | 5.4-6.2 | 5.4* | 78-97 | 95-107 | Pakistan |
|
|
588-700 | 24-32 | 81-113 | 83-113 | 121-139 | 99-117 | 21-29 | 4.7-5.4 | 5.5-6.6 | 3.0* | 60-86 | 89-110 | USA |
|
567-780 | 16-24 | 88-94 | 71-88 | 110-119 | 50-71 | 31-35 | 5.1-6.5 | 10.8-11.5 | – | 73-88 | 130-142 | USA |
|
|
as |
540-600 | 34-39 | 84-112 | 80-101 | 106-130 | 49-60 | 5-7 | 4-5 | 9–12 | – | 79-98 | 100-200 | USA |
|
|
736-800 | 27-32 | 123-142 | 104-115 | 147-160 | 112–128 | 23-38 | 4.6-5.9 | 6.1-6.9 | 6.3* | 81–91 | 103-120 | USA |
|
|
530-620 | 20-24 | 83-109 | 74-88 | 104-142 | 91-106 | 24-28 | 4.2-5.1 | 5.5-6.3 | 8.3* | 72-86 | 87-111 | Mexico |
|
|
484-578 | 21-25 | 88-105 | 69-96 | 79-115 | 78-95 | 21-27 | 4.3-5.2 | 5.5-6.8 | 3.4-4.3 | 81-95 | 99-125 | S. Africa | Malan et al. (2014) |
|
350–410 | 18-22 | – | – | – | 15-22 | – | – | – | – | – | – | USA |
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
550-676 | 19-23 | 103-122 | 86-101 | 125-141 | 86-108 | 25-32 | 3.9-4.9 | 5.4-7.5 | 8.7* | 80-90 | 99–133 | S. Africa |
|
|
332-499 | 17-23 | 74-113 | 58-87 | 96–130 | 44-70 | 18-27 | 3.4-4.2 | 6.5-8.7 | 3.7* | 71-96 | 110-230 | Egypt |
|
as |
495-570 | 19-32 | 97-116 | 87-98 | 110-131 | 91-125 | 19-26 | 4.4-5.4 | 4.0-6.5 | 6.7* | 78-110 | 81-111 | Mexico |
|
|
570-740 | 22-30 | 94-123 | 90-107 | 135-147 | 87-119 | 25 | 4.9 | 6.7 | – | 73-92 | 103-109 | N. Zealand |
|
as |
570-740 | 22-30 | 94-123 | 90-107 | 135-147 | 87-119 | 25 | 4.9 | 6.7 | – | 73-92 | 103-109 | N. Zealand |
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
Male | |||||
---|---|---|---|---|---|
Characters | Holotype | Paratypes | Hermaphrodite (1st Gen) paratypes | Female (2nd Gen) paratypes | Infective juvenile paratypes |
n | 1 | 20 | 20 | 20 | 25 |
Body length (L) | 760 | 769 ± 60 (652-863) | 3295 ± 286 (2907-4123) | 1366 ± 123 (1131-1608) | 544 ± 29 (496-591) |
a (L/BD) | 20.3 | 17 ± 1.5 (15-21) | 14.1 ± 1.1 (11.7-16.1) | 18 ± 1.4 (15-20) | 24 ± 3.0 (20-29) |
b (L/NL) | 7.8 | 8.1 ± 1.0 (5.8-9.7) | 23 ± 1.8 (21-27) | 11.4 ± 1.2 (9.0-13.6) | 4.7 ± 0.4 (4.1-5.4) |
c (L/T) | 26.2 | 31 ± 3.6 (23-36) | 42 ± 5.7 (34-51) | 20 ± 2.2 (16-24) | 8.2 ± 1.0 (7.6-8.6) |
c’ (T/ABW) | 1.1 | 1.4 ± 0.2 (0.6-1.7) | 2.2 ± 0.3 (1.7-2.6) | 2.8 ± 0.5 (1.9-3.6) | 4.6 ± 0.8 (3.4-5.8) |
V (VA/L × 100) | – | – | 48 ± 2.5 (45-55) | 48 ± 2.3 (41-51) | – |
Max. Body Width (MBD) | 37.5 | 44 ± 3.0 (40-51) | 233 ± 17 (209-274) | 77 ± 4.0 (68-83) | 23 ± 2.7 (18-27) |
Lip region width | 6.5 | 7.2 ± 0.8 (5.7-8.4) | 12.4 ± 0.8 (11.0-14.0) | 10.3 ± 0.9 (8.8-12.2) | – |
Stoma length | 9.5 | 11.1 ± 1.6 (8.7-13.9) | 14.9 ± 1.4 (13-18) | 13.6 ± 1.8 (10.4-16.0) | 13.8 ± 1.2 (12.1-16.0) |
Bulb length (BL) | 18.5 | 20 ± 1.8 (18-25) | 35 ± 3.6 (29-42) | 27 ± 2 (23-30) | 13.8 ± 1.8 (11.0-19.0) |
Pharynx length (PL) | 95.2 | 84 ± 7.1 (74-107) | 128 ± 6.3 (118-142) | 107 ± 6.9 (91-120) | 102 ± 7.0 (91-115) |
Nerve ring – ant. end (NR) | 68 | 63 ± 5.2 (56-74) | 93 ± 7.5 (78-108) | 81 ± 6.4 (69-97) | 55 ± 3.6 (52-64) |
Excretory pore– ant. end (EP) | 84.3 | 81 ± 10.1 (61-109) | 121 ± 11 (106-153) | 111 ± 10.8 (92-129) | 78 ± 3.4 (70-89) |
Neck length (Stoma+Pharynx, NL) | 98 | 96 ± 7.3 (84-117) | 143 ± 6.3 (134-159) | 120 ± 6.0 (107-132) | 115 ± 7.3 (103-131) |
Body width at neck base | 36 | 34 ± 1.9 (30-37) | 119 ± 8.9 (101-138) | 58 ± 4.3 (50-66) | 18 ± 3.0 (15-24) |
Vagina length | – | – | 28 ± 4.0 (20-38) | 19.2 ± 2.9 (15-26) | – |
Body width at vulva | – | – | 240 ± 21 (199-278) | 78 ± 3.8 (72-85) | – |
Vulva – ant. end (VA) | – | – | 1581 ± 151 (1369-1882) | 655 ± 47 (572-706) | – |
Vulva – post. End (PV) | – | – | 1713 ± 178 (1453-2241) | 710 ± 89 (559-949) | – |
Rectum length | – | – | 36 ± 4.6 (29-49) | 30 ± 3.8 (24-35) | 8.5 ± 1.9 (6.1-13.7) |
Anal body diam. (ABD) | 26.1 | 18 ± 2.4 (15-25) | 37 ± 5.5 (29-51) | 25 ± 4.5 (18-34) | 12.4 ± 1.8 (9.2-16.0) |
Tail with sheath length (T) | – | – | – | – | 56 ± 4.9 (49-64) |
Tail without sheath length | 29 | 25 ± 3.2 (21-29) | 80 ± 7.9 (63-98) | 68 ± 6.5 (62-88) | 30.4 ± 4.5 (22-39) |
Spicule length (SL) | 49 | 43 ± 4.1 (34-50) | – | – | – |
Gubernaculum length (GL) | 20.2 | 18 ± 1.5 (15-21) | – | – | – |
Stoma length/lip region width | 1.5 | – | 1.2 ± 0.2 (1.0-1.6) | 1.3 ± 0.1 (1.1-1.6) | – |
Nerve ring % (NR/NL × 100) | 69.4 | 67 ± 4.4 (58-75) | 65 ± 5.0 (56-78) | 67 ± 3.9 (61-75) | – |
Excretory pore % (EP/NL × 100) | 86 | 85 ± 8.5 (61-97) | 85 ± 8.3 (67-103) | 92 ± 7.6 (74-104) | – |
Rectum % (R/ABD × 100) | – | – | 90 ± 10 (80-130) | 128 ± 29 (90-181) | – |
D % (EP /PL × 100) | 88.5 | 96 ± 9.2 (69-111) | 95 ± 9.3 (74-114) | 104 ± 9.3 (82-118) | 78 ± 7.6 (66-98) |
E % (EP/T × 100) | 290 | 325 ± 49 (232-413) | 153 ± 24 (120-205) | 164 ± 23 (111-203) | 139 ± 13.4 (112-168) |
SW % (SL/ABD × 100) | 242 | 243 ± 47 (150-306) | – | – | – |
GS % (GL/SL × 100) | 41.2 | 42 ± 5.2 (35-57) | – | – | – |
H % (H/T × 100 | – | – | – | – | 46 ± 4.0 (37–55) |
Species | L | BD | EP | NR | NL | T |
|
|
|
|
V | ABD | D% | Country | Reference |
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
|
1279-2070 | 70-122 | 103-126 | 68-100 | 119-142 | 25-38 | – | – | – | 2.4* | 46-50 | 25-38 | – | Brazil | Andaló et al. (2006) |
|
1754-2628 | 86-129 | 154-182 | 79-119 | 129-167 | 80-108 | – | – | – | 3.8* | 43-49 | 24-33 | 100-113 | Chile |
|
|
3180-3850 | 160-220 | 174-214 | 93-118 | 155-183 | 71-93 | 21.4* | 18.8 | 41.5* | 3.1* | 42-53 | 22-31 | 114 | Australia |
|
1800-2400 | 100-162 | 122-162 | 83-102 | 108–145 | 40-65 | – | – | – | – | 41-50 | 23-40 | – | Argentina | Agüera de Doucet and Doucet (1986) | |
1690-3214 | 100-224 | 101-212 | 67-103 | 120-163 | 54-101 | – | – | – | 2.4* | 44-50 | 21-24 | 72-137 | Australia |
|
|
1513-2290 | 84-150 | 128–181 | 71-99 | 113–135 | 41-79 | 11-22 | 11-19 | 26-42 | 1.6–2.5 | 38-51 | 24-39 | 108–150 | India | Bhat et al. (2019a) | |
1226-1819 | 58-115 | 108-157 | 68-91 | 101-127 | 29-94 | 16-25 | 7.8-16 | 16-48 | 1.0-3.4 | 44-58 | 24-31 | 83-116 | India |
|
|
as |
2000-3500 | 90-180 | 105-240 | 88-140 | 162-200 | 75-108 | 12.5* | 7.8* | 31.2* | 2.0* | 42-48 | 33-35 | 100* | Argentina |
|
as |
2000-3300 | 184-240 | 146* | 126* | 148-177 | 71-93 | 11-15 | 14-21 | 26-46 | 2.8* | 48-53 | 33* | 95* | USA |
|
|
1335-2130 | 90-150 | 104-149 | 75-122 | 131-185 | 68-89 | 12-16 | 10-12 | 19-32 | – | 46-51 | 27-41 | – | Vietnam |
|
2060-2290 | 120-150 | 98-115 | 80-95 | 123-148 | 78-108 | 15-17 | 16-18 | 20-27 | - | 41–48 | 30-38 | 63-78 | India |
|
|
as |
2159-2666 | 117-194 | 143-156 | 90-112 | 128-144 | 41-80 | – | – | – | 2.9* | 36-51 | 21-35 | 104-111 | Thailand |
|
|
1581-3026 | 125-218 | 95-165 | 59-138 | 105-186 | 68-105 | 10-18 | 10–23 | 19–34 | 1.6-2.4 | 41-49 | 35–81 | 88-98 | China |
|
|
1231-2728 | 74–131 | 99-126 | 117-151 | 111–155 | 70-122 | – | – | – | 2.5* | 47-60 | 25-38 | – | Ireland |
|
|
1050-1420 | 56-84 | 69-106 | 69-94 | 106-125 | 56-78 | 17.5** | 14.4** | 22.2** | 3.1** | 44-51 | 19-27 | 78** | Egypt | Abd–Elgawad and Ameen (2005) |
|
2054-2548 | 120-156 | 110-168 | 86-122 | 126-178 | 69-87 | – | – | – | – | 44-50 | 32-42 | – | USA |
|
|
1640-2779 | 101-188 | 111-177 | 96–162 | 136-219 | 62-88 | – | – | – | 1.5* | 46-53 | 42* | – | USA |
|
|
600-1200 | 70-100 | 80-90 | 70-80 | – | – | – | – | – | – | 50-58** | – | – | Brazil |
|
|
2826-3983 | – | 148* | 161* | 219* | 28* | 13-19 | 12-21 | 47-67 | 0.8* | 43–49 | 33* | 92* | New Jersey |
|
|
1200-1800 | 76-113 | 118-138 | 88–96 | 120-139 | 66-88 | – | – | – | – | 40-53 | 22-32 | – | India |
|
1713-2242 | 110-156 | 135-172 | 77-92 | 120-138 | 61-83 | 13-17 | 11-18 | 22-36 | 1.9-2.9 | 44-50 | 27-33 | 102-128 | India |
|
|
1274-1993 | 70-135 | 105-129 | 84-111 | 124-155 | 64-83 | 12-18 | 10-13 | 16-31 | 2.6-4.9 | 45-52 | 22-30 | 77-99 | India | Bhat et al. (2021b) | |
as |
2100-2500 | 128-168 | 124-160 | 100-108 | 144-160 | 76-92 | – | – | – | – | 45-53 | 36-48 | – | China |
|
as |
1428-2533 | 71-161 | 108-157 | 73-141 | 120-182 | 66-95 | – | – | – | 3.3* | 43-55 | 22-38 | 74-112 | Australia | Plichta et al. (2009) |
as |
1300-2300 | 104-171 | 116-175 | 78-116 | 110-153 | 49-87 | – | – | – | – | 49-56 | 20-35 | – | USA |
|
as |
1413-1785 | 71-86 | 130-150 | 80-100 | 130-145 | 65-95 | 19-21 | 11-12 | 16-22 | 3.1* | 44-53 | 24-27 | 95* | Pakistan |
|
|
1600-2600 | 113-177 | 139-178 | 79-119 | 129–164 | 55-81 | – | – | – | 1.3* | 45-50 | 29-48 | 110* | USA |
|
– | – | – | – | – | – | – | – | – | – | – | – | – | USA |
|
|
as |
3500-4500 | 99-161 | 133-177 | 99-135 | 150-183 | 76–113 | 18* | 13* | 27* | 1.3* | 49-51 | 24-60 | 88* | USA |
|
|
1500-2500 | 95-140 | 158-206 | 105-120 | 155-168 | 70-101 | 15-19 | 10-16 | 18-32 | 2.6* | 47-51 | 25-38 | 119* | USA |
|
|
1144-2108 | 65-123 | 114-148 | 76-103 | 121-150 | 76-106 | – | – | – | – | 44-51 | 21-36 | – | Mexico |
|
|
1075-1697 | 76-129 | 102-125 | 73-90 | 115-132 | 63-75 | 13-17 | 9-14 | 17-24 | 2.3-3.1 | 40-53 | 22-32 | 83-104 | S. Africa | Malan et al. (2014) |
|
910-1520 | 62-80 | – | – | 152-172 | 86-105 | 11-14 | 50-51 | 10-11 | – | 38-50 | – | – | USA |
|
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1679-2937 | 102-229 | 151-196 | 87–139 | 148-180 | 55-111 | – | – | – | 1.3* | 45-50 | 25-72 | 97–120 | S. Africa |
|
|
830-1400 | 42-96 | 120-166 | 76-109 | 129-179 | 62-80 | – | – | – | 4.0* | 44-73 | 19-28 | 82* | Egypt |
|
as |
1500-2500 | 85-210 | 95-140 | 85-105 | 129-215 | 75-99 | – | – | – | 3.5* | 49-53 | 36-46 | 93* | Mexico |
|
|
– | – | – | – | – | – | – | – | – | – | – | – | – | N. Zealand |
|
as |
– | – | – | – | – | – | – | – | – | – | – | – | – | N. Zealand |
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|
Species | L | BD | EP | NR | NL | T |
|
|
|
|
V | ABD | D% | Country | Reference |
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
|
3517-5587 | 220-316 | 184-238 | 128-171 | 180-225 | 104-154 | – | – | – | 2.3* | 42-47 | 59-83 | 103* | Brazil | Andaló et al. (2006) |
|
1791-2904 | 88-122 | 165-206 | 101-132 | 174-200 | 72–112 | – | – | – | 2.7* | 39-48 | 30-46 | 90-114 | Chile |
|
|
3630-4390 | 160-180 | 189-217 | 121-130 | 189-205 | 81-93 | – | – | – | – | 41-47 | 40-53 | 106 | Australia |
|
4200-5600 | 175-242 | 163-225 | 125-152 | 180-220 | 50-87 | – | – | – | – | 35-45 | 45-62 | – | Argentina | Agüera de Doucet and Doucet (1986) | |
2686-4893 | 131-241 | 150-379 | 80–196 | 162–302 | 70-120 | – | – | – | 1.8* | 36-52 | 43-76 | 76-126 | Australia |
|
|
3086-5492 | 221-352 | 127-260 | 79-162 | 101–200 | 71-123 | 9.2-28 | 23-37 | 25-75 | 1.2-3.7 | 37-52 | 34-75 | 112-155 | India | Bhat et al. (2019a) | |
3916-5155 | 205-206 | 153-198 | 94-127 | 158-207 | 70-98 | 17-21 | 21-28 | 46-69 | 1.7-2.2 | 37-46 | 39-51 | 75-103 | India |
|
|
as |
5000-7500 | 250-575 | 250-340 | 132-196 | 235-300 | 100–140 | – | – | – | 1.8* | 40-50 | 70-120 | 102* | Argentina |
|
as |
3000-5100 | 200-344 | 250* | 250* | 163-286 | 76-100 | 11-18 | 11-25 | 30-63 | 2.2* | 45-52 | 62.5* | 80* | USA |
|
|
3135-4170 | 180-240 | 156-192 | 119-147 | 186-206 | 66-114 | 15-19 | 16-21 | 36-50 | 2.0* | 43-48 | 47-63 | 88* | Vietnam |
|
3250-3970 | 190-250 | 98-115 | 120-135 | 180-205 | 80-105 | 13-19 | 16-20 | 31-45 | – | 41-49 | 50-65 | 73-92 | India |
|
|
as |
2275-3952 | 108-183 | 156-214 | 118-144 | 158-193 | 56-87 | – | – | – | 2.3* | 41-56 | 30-53 | 86-113 | Thailand |
|
|
3671-5543 | 198-374 | 165-297 | 135-243 | 192-343 | 68-130 | 13-20 | 13-25 | 34-62 | 1.0-2.3 | 41-49 | 51-92 | 76-94 | China |
|
|
3030-5051 | 183-291 | 200-254 | 175-230 | 230-244 | 60-70 | – | – | – | 1.1* | 50-55 | 57-65 | 117* | Ireland |
|
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2100-3100 | 107-164 | 154-205 | 101-147 | 144-192 | 83-115 | – | – | – | 2.7* | 46-59 | 33-51 | 104* | Egypt | Abd–Elgawad and Ameen (2005) |
|
3731-5865 | 217-331 | 211-301 | 169-271 | 271-391 | 84-126 | – | – | – | 2.5* | 44-49 | 42-78 | 104* | USA |
|
|
3232-4928 | 157-267 | 200-277 | 143-217 | 132-271 | 65-96 | – | – | – | 1.2* | 44-55 | 42.6* | – | USA |
|
|
– | – | – | – | – | – | – | – | – | – | – | – | – | Brazil |
|
|
– | – | – | – | – | – | – | – | – | – | – | – | – | USA |
|
|
2300-3100 | 107-145 | 163-187 | 104-123 | 163-179 | 72-110 | – | – | – | – | 45-50 | 38-51 | – | India |
|
2751-4481 | 168-273 | 184-238 | 115-157 | 167-204 | 67-108 | 16-18 | 15-25 | 35-55 | 1.4-2.5 | 37-48 | 30-71 | 103-132 | India |
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2861-4227 | 152-208 | 140-179 | 119-146 | 165-186 | 79-114 | 16-23 | 17-24 | 30-47 | 1.5-2.4 | 39-55 | 37-56 | 81-100 | India | Bhat et al. (2021b) | |
as |
3550-5040 | 200-312 | 160-200 | 144-176 | 192-240 | 72-128 | – | – | – | – | 37-50 | 56-88 | 91* | China |
|
as |
2049-4288 | 93-209 | 103–288 | 82-210 | 146-317 | 90-196 | – | – | – | 2.4* | 40-48 | 40-80 | 90-147 | Australia | Plichta et al. (2009) |
as |
4000-7000 | 270-376 | 219-318 | 102-212 | 187-283 | 67–98 | – | – | – | – | – | 38-79 | – | USA |
|
as |
1939-4625 | 102-240 | 145-186 | 130-180 | 155-220 | 64–95 | 16–23 | 11–24 | 23–58 | 1.7* | 41-49 | 37-55 | 68-106 | Pakistan |
|
|
3000-4500 | 161–233 | 212-287 | 133-182 | 190-244 | 75-101 | – | – | – | 1.3* | 45-50 | 20-28 | 109* | USA |
|
– | – | – | – | – | – | – | – | – | – | – | – | – | USA |
|
|
as |
4000-5200 | 205-335 | 175-258 | 117-161 | 190-223 | 60-126 | – | – | – | 1.9* | 45-50 | 34-60 | – | USA |
|
|
2400-4900 | 120-133 | 193-270 | 139-178 | 106-269 | 95-124 | 14-24 | 12-21 | 23-49 | – | 45-50 | 36-86 | – | USA |
|
|
2440-4606 | 135-267 | 103-201 | 114-171 | 168-221 | 94-170 | – | – | – | 2.6* | 30-58 | 40-46 | 90* | Mexico |
|
|
2987-5498 | 168-289 | 152-209 | 112-152 | 166-220 | 79-120 | 14-23 | 18-28 | 37-58 | 1.7-3.4 | 39-47 | 26-56 | 77-112 | S. Africa | Malan et al. (2014) |
|
1350-2800 | 54-105 | – | – | – | 108-112 | – | – | – | – | – | – | – | USA |
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3373-4073 | 127-188 | 210-267 | 121-163 | 199-236 | 64-91 | – | – | – | – | 43-46 | 40-54 | 98-119 | S. Africa |
|
|
2200-2800 | 116-170 | 137-182 | 83–120 | 161-200 | 72-100 | – | – | – | – | 40-64 | 41-67 | – | Egypt |
|
as |
2856-5799 | 150-200 | 115-203 | 105-180 | 133-215 | 122-178 | – | – | – | 3.0* | 50-58 | 40-75 | - | Mexico |
|
|
– | – | – | – | – | – | – | – | – | – | – | – | – | N. Zealand |
|
as |
4000* | 247* | – | 181* | 236 | 90* | 16* | 17* | 44* | 1.7* | 46* | 53* | – | N. Zealand |
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Male | |||||
---|---|---|---|---|---|
Characters | Holotype | Paratypes | Hermaphrodite (1st Gen) paratypes | Female (2nd Gen) paratypes | Infective juvenile paratypes |
|
1 | 20 | 22 | 22 | 25 |
Body length (L) | 808.1 | 861 ± 29 (811-914) | 5127 ± 494 (4408-6179) | 2244 ± 203 (1954-2798) | 539 ± 21 (493-578) |
a (L/BD) | 19.0 | 18 ± 1.6 (15-22) | 16 ± 2.0 (13-20) | 12.3 ± 1.2 (10.5-15.0) | 22 ± 1.2 (19-24) |
b (L/NL) | 8.1 | 9.1 ± 1.1 (7.6-12) | 26 ± 4.3 (20-34) | 18 ± 1.8 (16-21) | 5.0 ± 0.4 (4.4-5.9) |
c (L/T) | 28.9 | 34 ± 4.2 (26-43) | 70 ± 10.4 (52-90) | 39 ± 7.4 (31-63) | 9.4 ± 0.6 (8.2-10.5) |
c’ (T/ABW) | 1.4 | 1.6 ± 0.3 (1.2-2.5) | 1.6 ± 0.3 (1.2-2.4) | 1.7 ± 0.2 (1.3-2.0) | 5.3 ± 0.6 (4.3-6.7) |
V (VA/L × 100) | – | – | 48 ± 4.3 (36-57) | 53 ± 4.2 (43-61) | – |
Max. Body Width (MBD) | 42.5 | 48 ± 3.6 (41-56) | 319 ± 41 (235-385) | 183 ± 23 (160-228) | 24 ± 0.9 (23-27) |
Lip region width | 6.2 | 7.4 ± 0.7 (6.2-8.8) | 11.7 ± 2.4 (9.2-19.2) | 10.1 ± 1.0 (7.7-11.4) | 4.0 ± 0.5 (3.2-5.2) |
Stoma length | 10 | 9.3 ± 1.0 (6.3-11) | 19 ± 2.0 (14-23) | 11.5 ± 1.7 (8.0-15.2) | 13.5 ± 1.0 (12.0-15.3) |
Bulb length (BL) | 20.2 | 22 ± 2.4 (19-28) | 40 ± 4.6 (28-49) | 30 ± 2.6 (28-38) | 20 ± 1.4 (17.1-23.0) |
Pharynx length (PL) | 95.2 | 86 ± 9.8 (57-100) | 182 ± 23 (155-211) | 113 ± 9.5 (101-133) | 95 ± 7.2 (82-111) |
Nerve ring – ant. end (NR) | 65.4 | 66 ± 5.3 (60-78) | 131 ± 22 (96-169) | 83 ± 7.3 (71-96) | 81 ± 6.3 (69-72) |
Excretory pore– ant. end (EP) | 96.2 | 93 ± 9.6 (77-109) | 150 ± 24 (108-190) | 113 ± 11 (100-133) | 89 ± 6.8 (72-99) |
Neck length (Stoma+Pharynx, NL) | 99.3 | 96 ± 9.6 (71-108) | 201 ± 21 (174-231) | 124 ± 10 (112-148) | 109 ± 6.9 (96-124) |
Body width at neck base | 34.5 | 36 ± 2.3 (31-40) | 167 ± 13 (133-188) | 95 ± 13.9 (74-121) | 23 ± 1.3 (19-26) |
Vagina length | – | – | 31 ± 4.0 (24-36) | 25 ± 6.4 (17-42) | – |
Body width at vulva | – | – | 331 ± 33 (257-379) | 185 ± 27 (153-230) | – |
Vulva – ant. end (VA) | – | – | 2470 ± 279 (1959-3038) | 1182 ± 129 (910-1397) | – |
Vulva – post. end (PV) | – | – | 2657 ± 279 (1990-3938) | 1062 ± 147 (860-1455) | – |
Rectum length | – | – | 36 ± 4.6 (30-41) | 27 ± 4.1 (19-39) | – |
Anal body diam. (ABD) | 19.6 | 17 ± 2.3 (13-22) | 47 ± 8.1 (34-58) | 35 ± 3.2 (31-41) | 11.1 ± 1.3 (8.6-14.1) |
Tail with sheath length (T) | – | – | – | – | 58 ± 3.1 (52-63) |
Tail without sheath length | 28 | 26 ± 3.3 (21-33) | 74 ± 8.3 (63-87) | 58 ± 8.2 (45-75) | 29.4 ± 2.5 (25-34) |
Spicule length (SL) | 54.1 | 45 ± 3.7 (38-55) | – | – | – |
Gubernaculum length (GL) | 18.7 | 20 ± 2.1 (15-25) | – | – | – |
Stoma length/lip region width | 1.6 | – | 1.6 ± 0.3 (1.1-2.1) | 1.2 ± 0.2 (0.8-1.7) | – |
Nerve ring % (NR/NL × 100) | 65.9 | 69 ± 9.9 (61-96) | 65 ± 9.4 (49-86) | 67 ± 4.8 (60-82) | – |
Excretory pore % (EP/NL × 100) | 96.9 | 98 ± 17 (78-134) | 75 ± 11 (51-95) | 67 ± 4.8 (60-82) | – |
Rectum % (R/ABD × 100) | – | – | 79 ± 17 (54-112) | 76 ± 13 (52-106) | – |
D % (EP /NL × 100) | 101.05 | 109 ± 21 (83-156) | 75 ± 11 (55-95) | 92 ± 7.9 (80-111) | 94 ± 12 (68-120) |
E % (EP/T × 100) | 343.6 | 365 ± 68 (236-503) | 206 ± 46 (145-303) | 197 ± 27 (145-246) | 154 ± 14 (128-184) |
SW % (SL/ABD × 100) | 276 | 270 ± 50 (170-320) | – | – | – |
GS % (GL/SL × 100) | 34.56 | 40 ± 10 (40-60) | – | – | – |
H % (H/T × 100 | – | – | – | – | 47 ± 5.6 (35-56) |