The paraleucobryoid Campylopus complex (Leucobryaceae) in South American páramos

The paraleucobryoid Campylopus complex consists of a small group of Andean species distinguished by a leaf cross section identical to that of Paraleucobryum. The complex differs from Paraleucobryum only in having cygneous rather than erect setae and capsule stomata absent rather than present. Here we treat four species, only two of which have previously been recognized in Campylopus. They are C. albidovirens Herzog, C. pittieri R.S.Williams, C. densifolius (Thér.) B.H.Allen & W.R.Buck, comb. nov. and C. ochyriorum B.H.Allen & W.R.Buck, sp. nov. A key is given to the four species; the two nomenclatural novelties are described in detail and illustrated.